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Gossypium tomentosum

Nutt. ex Seem. (1865)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Malvales Family: Malvaceae Genus: Gossypium

Hawaiian cotton, huluhulu, ma‘o [mao]

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Description

Key Characters:

Growth Form: Hawaiian cotton shrubs 0.5–1.5 m tall, softly white tomentose.

Stems:

Roots:

Leaves: Leaves simple. Alternate. Blades wider than long, usually 3–10(–12) cm wide, 3–lobed or some–times 5–lobed to about the middle, the lobes ovate, acute or subacuminate. Surfaces tomentose. Margins lobed but otherwise entire. Midvein on lower surface with basal gland absent or obscure. Petioles present. Stipules subulate, 0.5–0.8 cm long, caducous.

Flowers: Flowers solitary or few in short cymes, pedicel nectaries absent; involucral 3 bracts ovate, 15–25 mm long, dentate, with 5–9(–12) acute teeth. Flowers bisexual (perfect). Calyx erosely truncate, 7–9 mm long, sepals connate, the lobes valvate in bud. Corolla actinomorphic to moderately zygomorphic, corolla of 5 petals, bright yellow, without a maroon spot, fading greenish, 2–3.5(–4) cm long, campanulate to funnelform, the corolla of 5 petals, obovate, distinct from each other but adnate at base to staminal column, convolute in bud. Stamens 5 to numerous, monadelphous, forming a staminal column, staminal column included, 5-toothed at apex; antheriferous for most of its length or only in upper 1/2; anthers monothecal. Pollen globose, echinate. Ovary superior, 3-5-celled, the carpels borne in a single whorl or rarely seemingly superposed whorls, placentation axile; ovules several per cell; style exceeding the staminal column, unbranched, clavate; stigma 3-5-lobed, decurrent.

Fruit: Loculicidally dehiscent; usually glabrous capsules; 3–celled; coriaceous to woody; ovoid; cuspidate; 1.2–1.8 cm long; black–pitted. Seeds 2–4 per cell; ca. 5 mm long; densely covered with reddish brown lint; turbinate; with or without endosperm.

Ploidy: 2n = 52*

Habitat: Occurring primarily in arid; rocky; or clay coastal plains.

Elevation Range: 0–120 m.

Historical Distribution

Uses and Culture

USES

  • Flowers for yellow dye and leaves for green dye for kapa (Abbott 1993:66).

  • For treating the illness nahu‘aki o ka opu mai na makua mai ("gripping stimach ache"). To prepare, one needs to dry the flowers of ma‘o, nohu (Tribulus cistoides), ‘ilima (Sida fallax), and pua aloalo (cf. Hibiscus arnottianus) in the sun on niu (coconut, Cocos nucifera). The tap roots of ma‘o, ‘uhaloa (Waltheria indica), pōpolo (Solanum americanum), ‘ohi‘a ‘ai (Syzygyium malaccense), kōhonua‘ula (red sugarcane, Saccharum officinarum), and koali (Ipomoea spp.) are mashed and heated into a liquid form. The dried leaves are drunk with the liquid mixture (Chun 1994: 217–218).

  • Greenish (leaves), yellow (petals)

PROPAGATION/CULTIVATION

  • Gossypium tomentosum seeds are enclosed in a 3 part capsule. This oval woody fruit is 1/2 to 3/4 inch long and contains 6 to 12 seeds. The small seeds are covered with reddish brown fuzz.
  • Gossypium tomentosum grows easily from seed. Mew recommends scarification (penetrating the seed coat) to improve germination. Others recommend soaking the seeds in water for 24 hours to improve germination time. NTBG recommends cold water treatment; Bornhorst suggests hot water. Plant the seeds in a well-draining mix such as 2 parts perlite to 1 part sterile potting mix or a mix of 1 part perlite to 1 part peat moss. The pots should be placed in light shade and kept moist, but not over watered.
  • In his germination studies, Obata found that untreated seeds of Gossypium tomentosum had extrememly variable germination rates ranging from 1 to 75%. (Bornhorst 1996; Mew 1987; NTBG 1992; Obata 1967; Rauch 1997; Wagner 1990)
  • Gossypium tomentosum can be grown from cuttings. Criley reports success with both softwood and semi-hardwood terminal cuttings. He recommends using a rooting hormone of 2,000 parts per million (ppm) indolebutyric acid (IBA) in either a liquid or a talc dust form. Criley suggests either 1 part coarse perlite to 1 part vermiculite or 100% vermiculite as a rooting medium. His work was done using an intermittent mist system which was on for 6 to 8 seconds every 5 or 6 minutes; cuttings were rooted under 30% shade. Cuttings root in 3 to 4 weeks. (Criley 1998; Criley 1999)
  • Gossypium tomentosum can be grown from air layers. (Bornhorst 1996; Rauch 1997) [Data from Herring, E. C., & Criley, R. A. (2003). The Hawaiian Native Plant Propagation Web Site: Developing a Webbased Information Resource. HortTechnology, 13(3), 545-548. https://www.ctahr.hawaii.edu/hawnprop/]

Natural History

Statewide Status

Endemic

Island Status

Ni'ihau(Incl. Lehua) Endemic
Kuaihelani (Midway Atoll) Only found in cultivation
Kaua'i Unknown
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Kaho'olawe Endemic
Maui Endemic
Hawai'i Extirpated

Dispersal Agents


Pollinators

Notes

  • 0-120 m
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • also known as Ma'o, huluhulu

Bibliography

Name Published In: Fl. Vit.: 22 (1865)

Other References

Wagner et al. 1990:876 (Ni, K, O, Mo, L, M, Ka); Staples & Herbst 2005:386 (DESCR); Faccenda, Arthur, et al. 2025:193 (K† 1870, H† 1779); Naomi Worcester pers. comm. 2025 (Note: recently introduced to Ku for restoration and not yet established); Jonathan Plissner pers. comm. 2025 (Note: recently introduced to Mi for restoration and not yet established)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Gossypium tomentosum Manele Grade, stony ridge along Erosion zone. Preserved_Specimen Stone, B.C. 860 Lanai BISH 1955-07-02
2 Gossypium tomentosum Kaena Pt betw end of rd & Natural Area Reserve Coastal strand. Preserved_Specimen Shrub to 1m; flowers yellow. Annable, C.R. 3133 Oahu BISH 1996-11-13
3 Gossypium tomentosum Near Kolo, West Molokai Arid rocky plain. Preserved_Specimen Degener, O. 9591 Molokai BISH 1928-04-05
4 Gossypium tomentosum Puu Uala, flats near Dry flats. Preserved_Specimen Christensen, C. 173 Niihau BISH 1977-03-12
5 Gossypium tomentosum Near Kamakaipo, West Molokai Arid plain near ocean. Preserved_Specimen Degener, O. 9589 Molokai BISH 1928-05-16
6 Gossypium tomentosum In Diamond Head Crater Preserved_Specimen Shrub 2 m. high; flower yellow. Christophersen, E. 1316 Oahu BISH 1930-05-11
7 Gossypium tomentosum Near Alii Pond Arid clay plain. Preserved_Specimen Degener, O. 9592 Molokai BISH 1928-06-02
8 Gossypium tomentosum Palawai, top of Manele Grade Preserved_Specimen 1 m. shrub. St.John, H. 25556 Lanai BISH 1955-07-02
9 Gossypium tomentosum Nanakuli Seashore. Preserved_Specimen Wilder, G.P. Oahu BISH 1924-07-27
10 Gossypium tomentosum Manele Preserved_Specimen Munro, G.C. 120 Lanai BISH 1913-08-01
11 Gossypium tomentosum Lualualei, northwest of Nanakuli Beside road. Zone A of Ripperton and Hosaka. Preserved_Specimen Yellow-flowered sprawling shrub. Newhouse, W.J. Oahu BISH 1953-12-13
12 Gossypium tomentosum Kaena Pt. Preserved_Specimen shrub up to 3 m. Lv greyish, Fl lemon yellow, Fr [word illegible]. van Royen, P. 10240 Oahu BISH 1968-06-03
13 Gossypium tomentosum Waimanalo Arid region. Preserved_Specimen Bush, W. 9588 Oahu BISH 1929-05-26
14 Gossypium tomentosum Along route 72 between Koko Head and Wawamalu Beach Park Preserved_Specimen Common. Shrubby, 3 ft. tall; calyx pale green with black spots; corolla bright yellow; stamens yellow; style and stigma white. Crosby, M.R. 1544 Oahu BISH 1964-06-25
15 Gossypium tomentosum Waimea Arboretum and Botanical Garden Preserved_Specimen Tree form, 2 diameter trunk. Wilson, S. 35 Oahu BISH 2001-06-18
16 Gossypium tomentosum In Diamond Head Crater Preserved_Specimen Shrub ca. 2 m. height. Christophersen, E. 1432 Oahu BISH 1931-01-22
17 Gossypium tomentosum Kaluakoi Distr., Laau Point Preserved_Specimen Shrub with gray-green foliage, yellow flowers, brown linters on seed. Herbst, D.R. 5732 Molokai BISH 1976-03-09
18 Gossypium tomentosum Oahu Preserved_Specimen Colony of about 20 plants. Chock, A.K. 2051 Oahu BISH 1963-01-01
19 Gossypium tomentosum Kaluakoi Distr., Laau Point Preserved_Specimen Shrub with gray-green foliage, yellow flowers, brown linters on seed. Herbst, D.R. 5732 Molokai BISH 1976-03-09
20 Gossypium tomentosum In Diamond Head Crater Preserved_Specimen Shrub 2 m. high; flower yellow. Christophersen, E. 1361 Oahu BISH 1930-05-11
21 Gossypium tomentosum Kaohai Ahupuaa, In coastal kiawe forest just north of Awehi Road junction. Transect La, Station 5. Dry kiawe tree community with a[n] exotic and native shrub and mixed grass understsory. Preserved_Specimen Jacobi, J.D. 3007 Lanai BISH 1977-08-29
22 Gossypium tomentosum N Beach area, along road to Preserved_Specimen Decumbent shrub. Full flower and fruit. Flowers without noticeable fragrance. Gillett, G.W. 1940 Lanai BISH 1966-07-17
23 Gossypium tomentosum Bishop Museum, Atherton Halau Preserved_Specimen Prostrate shrub growing in full sun. Petal color mimosa. Stigmatic surface colored white. Lau, J. 1070 Oahu BISH 1985-06-10
24 Gossypium tomentosum Awalua Preserved_Specimen Kondo, Y. 1 Lanai BISH 1961-03-22
25 Gossypium tomentosum unknown Preserved_Specimen BISH
26 Gossypium tomentosum Manele Preserved_Specimen Munro, G.C. 29 Lanai BISH 1913-08-01
27 Gossypium tomentosum Waianae Mts., along road Preserved_Specimen Rock, J.F.C. 12825 Oahu BISH 1917-01-01
28 Gossypium tomentosum Along road 2 mi east of Kaena Point Preserved_Specimen A common shrub. Webster, G.L. 1093 Oahu BISH 1947-11-10
29 Gossypium tomentosum unknown Preserved_Specimen BISH
30 Gossypium tomentosum Manele Preserved_Specimen Munro, G.C. 28 Lanai BISH 1913-08-01
31 Gossypium tomentosum Manele Preserved_Specimen Munro, G.C. 22 Lanai BISH 1927-05-29
32 Gossypium tomentosum Queen's Beach off of Kalanianaole Hwy (Hwy 72) Preserved_Specimen Wagner, W.L. 4810 Oahu BISH 1983-06-04
33 Gossypium tomentosum Coco Head [Koko Head Preserved_Specimen Mann, H. 228 Oahu BISH 1864-01-01
34 Gossypium tomentosum Awalua Preserved_Specimen Kondo, Y. 1 Lanai BISH 1961-03-22
35 Gossypium tomentosum Ka Lae o Ka Laau Preserved_Specimen Forbes, C.N. 61 Molokai BISH 1912-06-01
36 Gossypium tomentosum W Maui, Pohakea Gulch Preserved_Specimen Degener, O. 17524 Maui BISH 1927-07-11
37 Gossypium tomentosum Waianae Mts., along road Preserved_Specimen Rock, J.F.C. 12825 Oahu BISH 1917-01-01
38 Gossypium tomentosum Manele, slopes above Preserved_Specimen Forbes, C.N. 286 Lanai BISH 1913-06-01
39 Gossypium tomentosum Coco Head [Koko Head Preserved_Specimen Mann, H. Oahu BISH 1864-01-01
40 Gossypium tomentosum Makapuu Point Preserved_Specimen Stokes, J.F.G. Oahu BISH 1920-05-02
41 Gossypium tomentosum Kapalama Heights, Kamehameha Girls' School Preserved_Specimen Judd Jr., A.F. Oahu BISH 1932-06-06
42 Gossypium tomentosum Kanoa Preserved_Specimen Swezey, O.H. Lanai BISH 1930-10-27
43 Gossypium tomentosum Along Koko Head-Waimanalo Road Preserved_Specimen Suehiro, A. Oahu BISH 1932-05-15
44 Gossypium tomentosum Molokai Preserved_Specimen Rock, J.F.C. Molokai BISH
45 Gossypium tomentosum Lualualei, Maili Point, Government road Preserved_Specimen Russ, G.W. Oahu BISH 1933-03-11
46 Gossypium tomentosum Maunalua Preserved_Specimen Judd Jr., A.F. Oahu BISH 1931-02-17
47 Gossypium tomentosum Kahoolawe Preserved_Specimen Forbes, C.N. Kahoolawe BISH 1913-02-10
48 Gossypium tomentosum Maunalei Gulch On basalt talus. Preserved_Specimen 1 m. clumps; flowers yellow, coma brown. St.John, H. 22645 Lanai BISH 1947-04-05
49 Gossypium tomentosum Along southwest shore between Barbers Point and Nanakuli, near base of Waianae Range Preserved_Specimen Skottsberg, C.J.F. 242210 Oahu BISH 1924-09-05
50 Gossypium tomentosum unknown Preserved_Specimen BISH
51 Gossypium tomentosum Waialae Nui Preserved_Specimen Forbes, C.N. Oahu BISH 1909-04-20
52 Gossypium tomentosum Near Koko Head Preserved_Specimen Degener, O. Oahu BISH 1930-01-01
53 Gossypium tomentosum Molokai Preserved_Specimen Stokes, J.F.G. Molokai BISH 1909-01-01
54 Gossypium tomentosum Kaunolu Preserved_Specimen Kondo, Y. Lanai BISH 1961-03-24
55 Gossypium tomentosum Makapuu Point Preserved_Specimen Stokes, J.F.G. Oahu BISH 1920-05-02
56 Gossypium tomentosum Koko Head Preserved_Specimen Rock, J.F.C. Oahu BISH
57 Gossypium tomentosum Molokai Preserved_Specimen Rock, J.F.C. Molokai BISH
58 Gossypium tomentosum unknown Preserved_Specimen Hillebrand, W.B. BISH
59 Gossypium tomentosum Honolulu, Foster Botanic Gardens Preserved_Specimen Krajina, V.J. 620131003 Oahu BISH 1962-01-31
60 Gossypium tomentosum Lanai Preserved_Specimen Forbes, C.N. 505 Lanai BISH 1917-09-01
61 Gossypium tomentosum In area 0.25 mi north of Smugglers' Cove The species is common in this area to Kaukaukapapa and extends from sea level to ca. 200 ft. elevation inland. Preserved_Specimen Mature plants badly chewed by goats. Char, W.P. 78.075 Kahoolawe BISH 1978-11-25
62 Gossypium tomentosum Central-western part of island, between Lua Kealialalo and coast In open Prosopis pallida scrub where ground caover dominated by Heteropogon contortus. Preserved_Specimen Low spreading shrub, 10' x 10' in area. Flowers sulfur yellow, closed in afternoon. Cuddihy, L.W. 371 Kahoolawe BISH 1980-04-22
63 Gossypium tomentosum Kaumalapau Highway, Koele/Keomuku Trail, along lower, N Lanai Very dry Prosopis forest. Preserved_Specimen Shrubs along road appearing dead with few leaves. Leaves alternate; orbicular, cleft into 3 lobes; base cordate. Capsule coriaceous, cuspidate, dehiscing into 3 valves. Spence, G. 307 Lanai BISH 1973-10-18
64 Gossypium tomentosum Kealakipapa Valley, Maunalua Edge of Prosopis forest. Preserved_Specimen Low spreading shrub; flowers light yellow. Fosberg, F.R. 10921 Oahu BISH 1935-06-27
65 Gossypium tomentosum East fork of Kawela Valley Arid plain. Preserved_Specimen Degener, O. 17522 Molokai BISH 1928-05-05
66 Gossypium tomentosum Kaumuhonu Bay, headland W Forming thicket on grassy headland. Preserved_Specimen Shrubs 1-2 m. tall; coma brown. St.John, H. 22782 Niihau BISH 1947-08-13
67 Gossypium tomentosum Beachfront at Kahe Point Among weedy grasses; population of scattered shrubs visible from the highway. Preserved_Specimen Takeuchi, W.N. 2441 Oahu BISH 1985-09-01
68 Gossypium tomentosum Manele Bay, arid slopes behind Common under Prosopis. Preserved_Specimen Sprawling shrub 1 m. ht. Involucral bracts stiff, prickly. Nagata, K.M. 2518 Lanai BISH 1982-08-27
69 Gossypium tomentosum Northeast of Koko Crater, Maunalua Rocky flats. Preserved_Specimen 5 ft. shrub, common. St.John, H. 10403 Oahu BISH 1930-02-28
70 Gossypium tomentosum Crater of Diamond Head, Honolulu Common in openings back from the margin of the lake bed. Preserved_Specimen 2-3 m. shrubs; flower lemon yellow. St.John, H. 17995 Oahu BISH 1937-05-28
71 Gossypium tomentosum W Maui, Lahaina District, Olowalu, SE of stream In Prosopis/Cenchrus Dry Forest. Preserved_Specimen Shrub; flowers yellow. Oppenheimer, H.L. 30602 Maui BISH 2006-03-11
72 Gossypium tomentosum West side of island, northeast of Lae o Kealaikahiki A frequent low shrub growing in partial shade under the canopy of Prosopis pallida scrub with a dense herbaceous cover of Heteropogon contortus. Preserved_Specimen Clarke, G. 359 Kahoolawe BISH 1980-04-21
73 Gossypium tomentosum Maunalei Gulch, mouth, near and along beach On sandy dry lands. Preserved_Specimen Quite common. Rock, J.F.C. 8037 Lanai BISH 1910-07-26
74 Gossypium tomentosum Maui Zoological & Botanical Gardens From Papawai. Point, near McGregor. Preserved_Specimen Davis, J. 51 Maui BISH 1978-07-03
75 Gossypium tomentosum Kaloko Along arid roadside. Preserved_Specimen Shrub with pale foliage, yellow flowers, and light brown fibers on seeds. Herbst, D.R. 751 Oahu BISH 1967-11-28
76 Gossypium tomentosum Kaali Cliffs, near base of Puu Uala Flat drylands. Preserved_Specimen Plants to 11 ft. tall. Christensen, C. 151 Niihau BISH 1977-02-12
77 Gossypium tomentosum Makapuu Head, Maunalua Growing in sandy soil in Prosopis grove. Preserved_Specimen Grant, M.L. 7065 Oahu BISH 1934-07-12
78 Gossypium tomentosum Hulopoe Bay, Palawai Dry rocky grassland. Preserved_Specimen 2 m. shrubs; flowers lemon yellow. St.John, H. 18777 Lanai BISH 1938-04-11
79 Gossypium tomentosum Shipwreck Beach, along road to beach, NE Lanai In dry forest area, now ravaged by grazing. With Nesoluma, Rauwolfia, Erythrina, Euphorbia, Sida. Preserved_Specimen Shrubby, to 3 ft., although mostly smaller. Flowers clear yellow; cotton white or sordid brownish. Carlquist, S. 2024 Lanai BISH 1966-07-17
80 Gossypium tomentosum W Maui, Pohakea Gulch Hot arid rocky plain. Preserved_Specimen Degener, O. 3650 Maui BISH 1927-07-11
81 Gossypium tomentosum Along road through Maalaea Village, 500 ft. from junction with Honoapiilani Hwy Associated with Desmanthus, grasses on banks of cane fields. Preserved_Specimen Low shrub. Ishikawa, S. 359 Maui BISH 1973-12-03
82 Gossypium tomentosum Manele Bay Arid sandy and clay shore. Preserved_Specimen Degener, O. 24232 Lanai BISH 1957-08-08
83 Gossypium tomentosum Between Kaloko and Makapuu Beach South side of ridge. along road. Preserved_Specimen Corn, C. Oahu BISH 1978-02-18
84 Gossypium tomentosum Roadside below Makapuu Point In arid, sunny coastal slopes, 0.25 mi. from ocean, with Prosopis chilensis. Preserved_Specimen Bushes 1 m. tall. Flowers brilliant yellow, 3.5 in. diam. Iltis, H.H. 300 Oahu BISH 1967-07-12
85 Gossypium tomentosum Maunalua near Makapuu Head Growing near roadside under Prosopis chilensis cover. Preserved_Specimen A shrub 3 ft. high; seeds covered with a brownish lint. Urata, U. Oahu BISH 1953-01-03
86 Gossypium tomentosum Roadside below Makapuu Point, 0.25 mi from ocean In arid, sunny coastal slopes, with Prosopis chilensis. Preserved_Specimen Bushes 1 m. tall. Flowers brilliant yellow, 3.5 in. diam. Iltis, H.H. 300 Oahu BISH 1967-07-12
87 Gossypium tomentosum Papawai Point, Ukumehame Rocky grassy slope, with Heteropogon. Preserved_Specimen 2 m. shrub; occasional. St.John, H. 25600 Maui BISH 1955-12-18
88 Gossypium tomentosum Highway 72, 0.5 mi west of Koko Head, near junction of road to Kuapa Pond, south Oahu Beside road, in grassy disturbed area now largely occupied by Leucaena leucocephala, although formerly an area of Erythrina forest. Preserved_Specimen Forming gray-green shrubs to 3 ft. tall. Flowers clear yellow; cotton of mature seeds tending to be brownish in color. Carlquist, S. 1685 Oahu BISH 1964-12-19
89 Gossypium tomentosum Kealakipapa Valley, Maunalua Edge of Prosopis forest. Preserved_Specimen Low spreading shrub; flowers light yellow. Fosberg, F.R. 10921 Oahu BISH 1935-06-27
90 Gossypium tomentosum Mahana Arid, rocky, clay plain. Preserved_Specimen Degener, O. 28579 Lanai BISH 1963-08-09
91 Gossypium tomentosum Roadside between Blowhole and Makapuu Point In rocky-sandy soil. Preserved_Specimen Spreading shrub ca. 1.5 m. high. Fujiwara, J. Oahu BISH 1960-10-04
92 Gossypium tomentosum Waianae Mts., rock rubble south of Puu Pueo Growing in a colony of Chamaesyce celastroides var. kaenana. Preserved_Specimen Shrub less than 4 ft. height but ca. 3 times as wide. Takeuchi, W.N. 2878 Oahu BISH 1986-09-28
93 Gossypium tomentosum Maunalua, between Koko Head and Makapuu Dry flats. Preserved_Specimen Bush 5 m. tall. Fosberg, F.R. 10358 Oahu BISH 1933-10-22
94 Gossypium tomentosum Nanakuli Sandy flat near shore. Preserved_Specimen 3 m. shrub. St.John, H. 25131 Oahu BISH 1953-12-13
95 Gossypium tomentosum Maunalua Rocky flat. Preserved_Specimen Flower yellow. St.John, H. 13189 Oahu BISH 1933-05-30
96 Gossypium tomentosum Diamond Head Beach Locally common in basin of crater; abundant in openings in algaroba; occasional below algaroba and Leucaena glauca. Preserved_Specimen Egler, F.E. 3745 Oahu BISH 1937-05-28
97 Gossypium tomentosum Between road and beach, a few hundres yards north of Smuggler's Cove Very dry conditions at time of collection. Preserved_Specimen Not flowering or fruiting and looking very weather worn. Corn, C. Kahoolawe BISH 1978-03-29
98 Gossypium tomentosum Makawao District, southwest coast near Hanakanaea; collected near helicopter LZ Abundant along coast and even vigorous in salt spray. Prosopis-Cenchrus Lowland Dry Forest; associated plants: Jacquemontia ovalifolia, Tragus berteronianus, Boerhavia repens, Macroptilium lathyroides, Sida fallax, Portulaca oleracea, Chenopodium murale, Atriplex semibaccata, Ipomoea cairica, Chamaesyce hirta, Abutilon incanum, Heteropogon contortus, Bothriochloa pertusa, Waltheria indica. Preserved_Specimen 1 m. tall shrub; in flower and fruit; corolla yellow. Wood, K.R. 1653 Kahoolawe BISH 1992-02-24
99 Gossypium tomentosum West side of island, northeast of Lae o Kealaikahiki A frequent low shrub growing in partial shade under the canopy of Prosopis pallida scrub with a dense herbaceous cover of Heteropogon contortus. Preserved_Specimen Clarke, G. 358 Kahoolawe BISH 1980-04-21
100 Gossypium tomentosum Maunalei Gulch, mouth, near and along beach On sandy dry lands. Preserved_Specimen Quite common. Rock, J.F.C. 8037 Lanai BISH 1910-07-26
101 Gossypium tomentosum Roadside (Kalanianaole Hwy) just before the (Koko Head side of) ascent to Makapuu Shrubs along road. Kiawe forest. Preserved_Specimen Yellow flowered. Stone, B.C. 2744 Oahu BISH 1959-03-29
102 Gossypium tomentosum Between Koko Crater and Makapuu Dry plain. Preserved_Specimen Hume, E.P. 184 Oahu BISH 1931-03-21
103 Gossypium tomentosum Between Makapuu Point and Hanauma Bay In grassy places along road. Preserved_Specimen Flowers lemon yellow; shrubs to 4 ft. Carlquist, S. 1804 Oahu BISH 1965-08-03
104 Gossypium tomentosum Near Makapuu Point Rocky slope, dry area. Preserved_Specimen Brown cotton. Baker, R.J. Oahu BISH 1932-09-18
105 Gossypium tomentosum Waianae Highway, on Kahe Point Rocky soil at roadside. Preserved_Specimen Bean, R.S. 317 Oahu BISH 1929-11-28
106 Gossypium tomentosum Honolulu, Diamond Head Xeric scrub along east and north upper slopes and rim. Occasional among Leucaena, floor of crater. Preserved_Specimen Shrub 2-2.5 or 3 m.; flowers bright yellow. Webster, G.L. 13831 Oahu BISH 1968-06-21
107 Gossypium tomentosum W Maui, Pohakea Gulch Hot arid rocky plain. Preserved_Specimen Degener, O. 3650 Maui BISH 1927-07-11
108 Gossypium tomentosum Oahu Rocky slope with Leucaena leucocephala. Preserved_Specimen One plant, several seedlings. Height 3 ft. Yellow flowers. Stephens, S.G. 2050 Oahu BISH 1963-03-10
109 Gossypium tomentosum 1 mi SW of Makapuu Point Rocky lowland. Preserved_Specimen Dunn, J. Oahu BISH 1935-04-13
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